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A 3-year postdoctoral position is available to work on a project to study inter-genomic conflict in gynodioecy and its effects on molecular evolution of mitochondrial genomes in plants in the genus Plantago. The position will be based in Deborah Charlesworth's group, in the Institute of Evolutionary Biology, University of Edinburgh, collaborating with Kirsten Wolff's group at Newcastle University.

This project will use molecular evolutionary approaches to investigate the maintenance of cytonuclear polymorphisms in gynodioecious plants in the genus Plantago, to test whether an 'arms race' situation prevails, versus long-term balancing selection. The Edinburgh part of the work will include obtaining and analysing DNA sequences of multiple nuclear and mitochondrial genes, to compare diversity, and test for long-term balancing selection. Non-gynodioecious species will be studied, as well as gynodioecious ones, to provide outgroups for sequence analyses and to test whether the high diversity expected under long-term balancing selection is confined to the gynodioecious species. The project also includes testing for paternal inheritance/leakage by testing for heteroplasmy in natural populations of P. lanceolata, using analyses of mitochondrial DNA sequence haplotypes and studies of offspring of reciprocal crosses, and modelling, to assess the possibility that within-species polymorphism could explain the highly elevated mutation rates estimated in Plantago lineages.

Fixed Term: 3 years

Salary Scale: £30,122 - £35,938

Please Quote Ref: 3016025

Closing Date: 20 August 2012

For further particulars https://www.jobs.ed.ac.uk/jobs/index.cfm?action=jobdet&jobid=3016025 and an application pack visit our website (www.jobs.ed.ac.uk) or telephone the recruitment line on 0131 650 2511.